Bryophytes are the group of plants that are the closest extant relative of early terrestrial vascular plants. They comprise three separate evolutionary lineages, which are today recognized as liverworts (phylum Marchantiophyta), mosses (phylum Bryophyta), and hornworts (phylum Anthocerotophyta). Liverworts are viewed as the plants most closely related to the ancestor that moved to land. Bryophytes play an important role in earth ecology by contributing to water retention initiation of soil formation and nutrition decomposition. The project aims on our better understanding of genome architecture, chromosome structure and epigenetic pathays in these early diverging plants.