(1) above the reef slope just below the water surface (surface water); (2) 15 m away from the reef slope at a water depth of 15 m (open ocean water); above the reef slope, 1 m above the seabed at water depths of 5 m (3) and 15 m (4) (intermediate water); directly above BCMs (within 1 cm of the seabed) at water depths of 5 m (5) and 15 m (6) (bottom water), and directly above BCM-free substrate at water depths of 5 m (7) and 15 m (8) (control bottom water).